In the new film, "X-Men: Apocalypse," the titular X-villain unleashes his Four Horsemen -- mutants who serve his cause, including some very familiar faces (Magneto, Psylocke, Storm and Angel). Nearly as long as Apocalypse has been appearing in Marvel's comic books, he has been aided by his minions. G based on the Biblical Four Horsemen of the Apocalypse (Pestilence, Famine, War and Death), the group has had many incarnations over the years.

RELATED: En Sabah Who? The History of the Villain of "X-Men: Apocalypse"

Comprise of outright villains to X-Men themselves, the Four Horsemen have plagued the X-Men for decades, transforming their powers and psyches but always remaining a terrifying force for evil. With their on-screen debut having arrived, we look at the many Horsemen Apocalypse has assembled over the years, from the original group that debuted in 1986, to the current foursome wreaking havoc in Marvel Comics' "Apocalypse Wars" mini-event.

The Original Four Horsemen



Apocalypse officially debuted in "X-Factor" #6 (following a brief silhouetted cameo in the previous issue). He didn't have Horsemen at the time, but it wasn't long before he began to assemble them. In "X-Factor" #10 (by Louise Simonson, Walter Simonson and Bob Wiacek), during the middle of the Mutant Massacre, Apocalypse saved one of the Morlocks, Plague, and recruited her as his new Pestilence. In the following issue, Apocalypse recruited a former soldier as his War, and the next issue he recruited an anorexic girl as Famine. The three debuted as his Horsemen officially in "X-Factor" #15, but they didn't find their fourth until "X-Factor" #24. In a shocking move, founding X-Man and former X-Factor member Warren Worthington III (AKA Angel) joined the group as Death, complete with new artificial wings and calling himself Archangel. Worthington eventually broke free from the group, and Pestilence was killed in battle.

X-Cutioner's Song



At the start of "X-Cutioner's Song," the two surviving original members of the Four Horsemen, along with a new Death, the Morlock known as Caliban, attacked Jean Grey and Scott Summers in "Uncanny X-Men" #294 (by Scott Lobdell, Brandon Peterson and Terry Austin). They believed they were serving Apocalypse, but in reality they were following the orders of Mister Sinister, who impersonated Apocalypse to get them to work for him.

World War Hulk



In a brief tangential story in "Incredible Hulk" #456 (by Peter David, Joe Kubert and Mark Farmer), Apocalypse manipulated the Hulk into being his new War for a brief period of time.

The Twelve



During the lead-up to the X-Men storyline, "The Twelve," the X-Men were attacked by Apocalypse's new Death. In "Astonishing X-Men" #3 (by Howard Mackie, Brandon Peterson and Tim Townsend), Death actually managed to kill Wolverine despite his mutant healing factor. However, that Wolverine turned out to be a Skrull. So where was the real Wolverine? Well, in "X-Men" #95 (by Alan Davis, Tom Raney and Scott Hanna), it was revealed that Apocalypse had indeed recruited and brainwashed Wolverine into becoming his new Death, restoring the mutant's Adamantium-laced bones in the process. (The metal had been pulled form his body by Magneto in an earlier storyline.) In "X-Men" #97 (by Alan Davis, Terry Kavanagh and Mark Farmer), the rest of the Four Horsemen were revealed, with Deathbird as War, Caliban as Pestilence and Ahab as Famine. They were quickly defeated.

The Blood of Apocalypse



In "X-Men" #182 (by Peter Milligan and Salvador Larroca), Apocalypse began recruiting his newest Horsemen, with the mutant known as Gazer battling to the death against another unnamed mutant to earn the right of becoming the new War. Sunfire, at the time, had lost his legs to Lady Deathstrike and his powers to Rogue. So, naturally, Apocalypse recruited him as his new Famine. In "X-Men" #183 and #184, Gambit actually volunteered to be the new Death in an attempt to keep an eye on Apocalypse (which turned out not to be the brightest idea Gambit ever had). In "X-Men" #185, readers learned Polaris had become Apocalypse's new Pestilence. The strangest thing about this storyline is that these Four Horsemen never actually appear together. War is killed in "X-Men" #186, which ended the storyline leaving Polaris returning to normal and Gambit and Sunfire heading off together.

The Apocalypse Solution



In "Uncanny X-Force" #3 (by Rick Remender and Jerome Opeña), it was revealed that Apocalypse and his not-so-trusty servant, Ozymandias, had been collecting special people throughout history, to be put aside and transformed into Horsemen in the case of extreme emergency. When Apocalypse was resurrected as a young boy, this was deemed such a time. These so-called "Final Horsemen" included War, a Roman mutant people believed was the mythological Minotaur; Famine, a Confederate soldier during America's Civil War; Pestilence, a 19th Century Japanese geisha who unleashed a swarm of beetles on the women of her village; and Death, a 9th Century Persian with the ability to kill people by giving them various ailments.

After the young Apocalypse was killed, Archangel slowly transformed into his successor. During another battle with X-Force, Death was killed. Psylocke then became Archangel's new Death in "Uncanny X-Force" #17, albeit very briefly.

The Apocalypse Twins



After Archangel was dealt with, the next Apocalyptic threat occurred when the Apocalypse Twins were introduced. The twins were born of Archangel and his Pestilence, and were taken into the future by Kang. They then returned, grown and ready to take on the Uncanny Avengers. Their horsemen were all Death, created by resurrecting four dead heroes and villains -- Grim Reaper, Daken (Wolverine's son), Banshee and the Sentry. The Horsemen eventually broke free of the Apocalypse Twins' control and the Avengers Unity Squad was able to defeat the Twins' evil plan.

Apocalypse Wars



In the current "Apocalypse Wars" storyline running in "Extraordinary X-Men," Colossus endes up in the future with a few of his young charges. When the X-Men travel to the future after them in "Extraordinary X-Men" #8 (by Jeff Lemire, Humberto Ramos and Victor Olazaba), the students have now grown. Meanwhile, at the end of the issue, we see Colossus has joined up with the Four Horsemen, along with a female Moon Knight, future Deadpool and some sort of Venom symbiote. Time will tell what will happen to this group.

Retroactive Four Horsemen



In "Uncanny Avengers" #6 (by Rick Remender and Daniel Acuña), readers witnessed Thor battle against Apocalypse and perhaps the earliest version of the Four Horsemen in the 11th Century. Little is known about this group.

Age of Apocalypse



There have been far too many alternate reality Four Horsemen to detail them all, and the "Age of Apocalypse" Four Horsemen are the ones most worthy of calling out. By the time the crossover/reality began, ten years in real time after Apocalypse first declared war on humanity, En Sabah Nur had stopped differentiating between the Horsemen. In this reality they were comprised of Holocaust, Mister Sinister, Mikhail Rasputin and Lord Abyss, and while this lineup has not been seen since, their impact was felt in helping establish the threat of Apocalypse as one of the biggest in the Marvel Universe.

Which grouping represents your favorite Four Horsemen? Let us know in the comments!